Its not current in the way we think of it. Think of it more as load balance between the plates and the speaker, instead of a pure resistive load. You know, there is no "impedance" of a transformer, only its ratio
If you take a tube amp, using an output transformer, and run that transformer into nothing... an open circuit... it will self destruct sooner than later, and depending on the transformer MUCH sooner.... If you take a tube amp, with an output transfomer and run it into a dead short, nothing happens. I know.... its a leap of faith, just go with it. The EXACT OPPOSITE is true for capacitor coupled amps (rare) or for amps with silicon solid state output. You can run those higher impedance loads all day long, but not lower. Back to tube amps - Marshall, early classic ones with "Radio Spares" transformers are notorious for burning up when you mis-set the impedance plug. Part of their tone, is the lack of headroom, the ability to turn electricity into magnetism, in the output transformer. They also had light windings... the mismatch of impedance, plus little in the way of overkill... and poof. Its also interesting to note, that the poof occurs almost always on the primary side, not the speaker side. Makes ya go hmmm. Fender uses a switched jack, such that if you dont have a speaker plugged in - it goes direct short - hot to chassis ground. And they'll sit there all day and night, for about forever, without burning up the transformer. If you lose a speaker cable, or blow the voice coil on a single speaker amp... it will take a while, but it will go poof in a Fender. Its not nearly the fast blow a Marshall has, because Leo very generously oversized his transformer. A Fender, or at least the classic ones, will generally handle a 2x mismatch just fine. More than that, you're in more dangerous territory... Going lower, no problem though.